Melanism in the jaguar is conferred by a dominant allele, and in the leopard (Panthera pardus) by a recessive allele. Close examination of the color of these black cats will show that the typical markings are still present, but are hidden by the excess black pigment melanin, giving an effect similar to that of printed silk. This is called "ghost striping". Melanistic and non-melanistic animals can be littermates. It is thought that melanism may confer a selective advantage under certain conditions since it is more common in regions of dense forest, where light levels are lower.