Another benefit is gaining quite a bit more followers. I have not done any study, but I am pretty sure that commenting gains me a ton more followers than posting articles on my own blog. I also tend to find more people that I want to follow. I'm not sure there is any direct payout effect from followers, but they do seem to add to the conversation.
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I would tend to agree with you @diggndeeper.com
There is no doubt commenting gets on recognized...also by some bigger fish. A bran new person most likely isnt going to interact with a dolphin on the new person's post. However, when one comments, the chances of interacting with someone like that goes up greatly (and gaining an upvote and follower).